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The dog (Canis lupus familiaris) is a domestic subspecies of the wolf, a mammal of the Canidae family of the order Carnivora. The term encompasses both feral and pet varieties and is also sometimes used to describe wild canids of other subspecies or species. The domestic dog has been (and continues to be) one of the most widely-kept working and companion animals in human history, as well as being a food source in some cultures. The dog is also the first animal from Earth to enter into space and fly into orbit.

The dog has developed into hundreds of varied breeds. Height measured to the withers ranges from a few inches in the Chihuahua to a few feet in the Irish Wolfhound; color varies from white through grays (usually called blue) to black, and browns from light (tan) to dark ("red" or "chocolate") in a wide variation of patterns; and, coats can be very short to several centimeters long, from coarse hair to something akin to wool, straight or curly, or smooth.

Contents
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* 1 Etymology and taxonomy
* 2 Origin and evolution
o 2.1 Origins
o 2.2 Ancestry and history of domestication
+ 2.2.1 Wolf ancestors
+ 2.2.2 Rate of domestication
o 2.3 Development of dog breeds
+ 2.3.1 Neoteny in the rapid evolution of diverse dog breeds
+ 2.3.2 Breed popularity
* 3 Physical characteristics
o 3.1 Sight
o 3.2 Hearing
o 3.3 Smell
o 3.4 Coat color
o 3.5 Sprint metabolism
* 4 Behavior and Intelligence
o 4.1 Intelligence
+ 4.1.1 Evaluation of a dog's intelligence
o 4.2 Human relationships
o 4.3 Dog communication
o 4.4 Laughter in dogs
* 5 Reproduction
o 5.1 Life cycle
o 5.2 Spaying and neutering
o 5.3 Overpopulation
* 6 Working, utility and assistance dogs
* 7 Show and sport (competition) dogs
* 8 Dog health
o 8.1 Morbidity (Illness)
+ 8.1.1 Diseases
+ 8.1.2 Parasites
+ 8.1.3 Common physical disorders
o 8.2 Mortality (Lifespan and causes of death)
o 8.3 Diet
+ 8.3.1 Nutrition
+ 8.3.2 Dangerous substances
o 8.4 Dog abuse
+ 8.4.1 Signs of dog abuse
* 9 Dog meat for human consumption
* 10 Dogs in religion
o 10.1 Chinese tradition
o 10.2 Christianity
o 10.3 Islam
o 10.4 Atheism and criticism of religion
* 11 See also
* 12 References
* 13 Further reading
* 14 External links

Etymology and taxonomy

The English word dog, in common usage, refers to the domestic pet dog, Canis lupus familiaris. The species was originally classified as Canis familiaris by Linnaeus in 1758. In 1993, dogs were reclassified as a subspecies of the gray wolf, Canis lupus, by the Smithsonian Institution and the American Society of Mammalogists. "Dog" is sometimes used to refer collectively to any mammal belonging to the family Canidae (as in "the dog family"), such as wolves, foxes, and coyotes. Some members of the family have "dog" in their common names, such as the African hunting dog.

The English word dog might derive from the Old English docga, a "powerful breed of canine". The English word hound is a cognate of German Hund, Dutch hond, common Scandinavian hund, Icelandic hundur which, though referring to a specific breed in English, means "dog" in general in the other Germanic languages. Hound itself derives from the Proto-Indo-European *kwon-, which is the direct root of the Greek κυων (kuōn) and the indirect root of the Latin canis through the variant form *kani-.[1]

In breeding circles, a male canine is referred to as a dog, while a female canine is called a bitch. The father of a litter is called the sire, and the mother of a litter is called the dam. Offspring are generally called pups or puppies until they are about a year old. A group of offspring is a litter. The process of birth is whelping. Many terms are used for dogs that are not purebred.

Origin and evolution

Origins

Main article: Origin of the domestic dog

Some research appears to show that dogs were domesticated from wolves as recently as 15,000 years ago,[2] or perhaps as early as 100,000 years ago based upon recent genetic, fossil and DNA evidence.[3] Other research suggests that dogs have only been domesticated for a much shorter amount of time and were domesticated from populations of wild dogs, which had previously diverged from wolves.[4][5]

New evidence suggests that dogs were first domesticated in East Asia, possibly China,[6] and the first peoples to enter North America took dogs with them from Asia. Genetic research has identified 14 ancient dog breeds, with the oldest being the Chow Chow, Shar Pei, Akita Inu, Shiba Inu and Basenji. Because many of the 14 breeds are associated with China and Japan, the theory that the dog originated in Asia seems to be likely.[6]

As humans migrated around the planet a variety of dog forms migrated with them. The agricultural revolution and subsequent urban revolution led to an increase in the dog population and a demand for specialization. These circumstances would provide the opportunity for selective breeding to create specialized working dogs and pets.

Ancestry and history of domestication
It has been suggested that this article or section be merged into Origin of the domestic dog. (Discuss)

Main article: Origin of the domestic dog

This ancient mosaic, likely Roman, shows a large dog with a collar hunting a lion.
This ancient mosaic, likely Roman, shows a large dog with a collar hunting a lion.

Molecular systematics indicate that the domestic dog (Canis lupus familiaris) descends from one or more populations of wild wolves (Canis lupus). As reflected in the nomenclature, dogs are descended from the wolf and are able to interbreed with wolves.

The relationship between human and canine has deep roots. Wolf remains have been found in association with hominid remains dating from 400,000 years ago. Converging archaeological and genetic evidence indicate a time of domestication in the late Upper Paleolithic close to the Pleistocene/Holocene boundary, between 17,000 and 14,000 years ago. Fossil bone morphologies and genetic analysis of current and ancient dog and wolf populations have not yet been able to conclusively determine whether all dogs descend from a single domestication event, or whether dogs were domesticated independently in more than one location. Domesticated dogs may have interbred with local populations of wild wolves on several occasions (so-called introgression).

The earliest dog fossils, two crania from Russia and a mandible from Germany, date from 13,000 to 17,000 years ago. Their likely ancestor is the large northern Holarctic wolf, Canis lupus lupus. Remains of smaller dogs from Mesolithic (Natufian) cave deposits in the Middle East, dated to around 12,000 years ago, have been interpreted as descendants of a lighter Southwest Asian wolf, Canis lupus arabs. Rock art and skeletal remains indicate that by 14,000 years ago, dogs were present from North Africa across Eurasia to North America. Dog burials at the Mesolithic cemetery of Svaerdborg in Denmark suggest that in ancient Europe dogs were valued companions.

Genetic analyses have so far yielded divergent results. Vilà, Savolainen, and colleagues (1997) concluded that dogs split off from wolves between 75,000 and 135,000 years ago, while a subsequent analysis by Savolainen et al. (2002) indicated a "common origin from a single gene pool for all dog populations" between 40,000 and 15,000 years ago in East Asia. Verginelli et al. (2005), however, suggest both sets of dates must be reevaluated in light of recent findings showing that poorly calibrated molecular clocks have systematically overestimated the age of geologically recent events. On balance, and in agreement with the archaeological evidence, 15,000 years ago is the most likely time for the wolf-dog divergence.

Verginelli examined ancient DNA evidence from five prehistoric Italian canids carbon-dated to between 15,000 and 3,000 years old, 341 wolves from several populations worldwide, and 547 purebred dogs. Their results indicate multiple independent origins of dogs and/or of frequent interbreeding between early proto-dogs and wolves throughout a vast geographic range. The detailed history remains unexplored and until further evidence is available, the following section on wolf ancestors must be considered purely speculative.

Wolf ancestors
Some breeds of dog retain a primitive, wolf-like appearance to this day
Some breeds of dog retain a primitive, wolf-like appearance to this day

Although all wolves belong to the species Canis lupus, there are (or were) many subspecies that had developed a distinctive appearance, social structure, and other traits. For example, the Japanese Wolf and the Eastern Timber Wolf possess different distinctive coloration, hunting and social structures. The origin of the dog is so ancient and so worldwide that many varieties of wolf played a part in it. It is wrong to say that dogs descended from modern wolves. They descended from ancestral wolves, and this difference must always be kept in mind. Ancestral wolves of many varieties existed all over the world.

The Indian Wolf is thought to have contributed to the development of more breeds of dogs than other subspecies. Many of today's wild dogs, such as the dingo and the pariah dogs, are descended from this wolf. The Indian Wolf is also thought to have bred with descendants of the European wolf to create the Mastiffs and eventually leading to the development of such diverse breeds as the Pug, the Saint Bernard, and the Bloodhound. The Tibetan Mastiff is an example of an ancient breed.

The European wolf, in turn, may have contributed many of its attributes to the Spitz dog types, most terriers, and many of today's sheepdogs. The Chinese wolf is probably ancestor to the Pekingese and toy spaniels, although it is also probable that descendants of the Chinese and European wolves encountered each other over the millennia, contributing to many of the oriental toy breeds.

The Eastern Timber Wolf is a direct ancestor to most, if not all, of the North American northern sled dog types. This interbreeding still occurs with dogs living in the Arctic region, where the attributes of the wolf that enable survival in a hostile environment are valued by humans. Additionally, unintentional crossbreeding occurs simply because dogs and wolves live in the same environment. The general reproductive isolation which is required to define dogs and wolves as separate species is purely a result of lack of opportunity, stemming from a general mutual unfamiliarity, suspicion, mistrust, and fear.

The phenotypic characteristics that distinguish a wolf from a dog are tenuous. Wolves typically have a "brush tail" and erect ears. While some dog breeds possess one of these characteristics, they rarely possess both.

Rate of domestication

Current research indicates that domestication, or the attributes of a domesticated animal, can occur much more quickly,[7] [not in citation given] than previously believed. Domestication of a wild dog may occur within one or two human generations with deliberate selective breeding. It is also now generally believed that initial domestication was through mutual desire. Wild canines who scavenged around human habitations received more food than their more skittish or fearful counterparts. Canines who attacked people or their children were likely killed or driven away, while those more friendly animals survived. Canines would have been beneficial by chasing away other vermin or scavengers. With their sharp senses, they would also be valuable as an alarm against marauding predators. The relationship is theorized to have developed in this way.

A 2007 Animal Planet documentary discussed a 1900-1920 domestication experiment in Russia that led to rapid unusual changes in the coats of foxes.

Development of dog breeds
Dogs have been bred into an amazing variety of shapes, colors and sizes. Variation can be wide even within a breed, as with these Cavalier King Charles Spaniels.
Dogs have been bred into an amazing variety of shapes, colors and sizes. Variation can be wide even within a breed, as with these Cavalier King Charles Spaniels.

There are numerous dog breeds, with over 800 being recognized by various kennel clubs worldwide. Many dogs, especially outside the United States of America and Western Europe, belong to no recognized breed A few basic breed types have evolved gradually during the domesticated dog's relationship with humans over the last 10,000 or more years, but most modern breeds are of relatively recent derivation. Many of these are the product of a deliberate process of artificial selection. Because of this, some breeds are highly specialized, and there is extraordinary morphological diversity across different breeds. Despite these differences, dogs are able to distinguish dogs from other kinds of animal.

The definition of a dog breed is a matter of some controversy. Depending on the size of the original founding population, closed gene pool breeds can have problems with inbreeding, specifically due to founder effect. Dog breeders are increasingly aware of the importance of population genetics and of maintaining diverse gene pools. Health testing and new DNA tests can help avoid problems, by providing a replacement for natural selection. Without selection, inbreeding and closed gene pools can increase the risk of severe health or behavioral problems. Some organizations define a breed more loosely, such that an individual may be considered of one breed as long as 75% of its parentage is of that breed. These considerations affect both pets and the show dogs entered in dog shows. Even prize-winning purebred dogs sometimes possess crippling genetic defects due to founder effect or inbreeding.[8] These problems are not limited to purebred dogs and can affect cross-breed populations.[9] The behavior and appearance of a dog of a particular breed can be predicted fairly accurately, while mixed-breed dogs show a broader range of innovative appearance and behavior.

In February 2004, the Canine Studies Institute in Aurora, Ohio, arranged recognized breeds of dogs into ten categories.
This puppy is a mix of many breeds.
This puppy is a mix of many breeds.

Mixed-breed dogs or Mongrels are dogs that do not belong to specific breeds, being mixtures more than two in variant percentages. Mixed breeds, or dogs with no purebred ancestry, are not inherently better or worse than purebred dogs as companions, pets, working dogs, or competitors in dog sports. Sometimes mixed-breed dogs are deliberately bred, for example, the Cockapoo, a mixture of Cocker Spaniel and Miniature Poodle. These are known as cross-breeds. Such deliberate crosses may display some degree of hybrid vigor and other desirable traits, but may or may not inherit any of the desired traits of their parents, such as temperament or a particular color or coat. Without genetic testing of the parents, the crosses can end up inheriting genetic defects that occur in both parental breeds. Deliberately crossing two or more breeds is also a manner of establishing new breeds.

Neoteny in the rapid evolution of diverse dog breeds
It has been suggested that this article or section be merged into Origin of the domestic dog. (Discuss)

This rapid evolution of dogs from wolves is an example of neoteny or paedomorphism. As with many species, the young wolves are more social and less dominant than adults; therefore, the selection for these characteristics, whether deliberate or inadvertent, is more likely to result in a simple retention of juvenile characteristics into adulthood than to generate a complex of independent new changes in behavior. (This is true of many domesticated animals, including humans themselves, who have many characteristics similar to young bonobos.) This paedomorphic selection naturally results in a retention of juvenile physical characteristics as well. Compared to wolves, many adult dog breeds retain such juvenile characteristics as soft fuzzy fur, round torsos, large heads and eyes, ears that hang down rather than stand erect, etc.; characteristics which are shared by most juvenile mammals, and therefore generally elicit some degree of protective and nurturing behavior cross-species from most adult mammals, including humans, who term such characteristics "cute" or "appealing".

The example of canine neoteny goes even further, in that the various breeds are differently neotenized according to the type of behavior that was selected.[10]

* Livestock guardian dogs exhibit the controlled characteristics of hunting dogs. Members of this group, such as Border Collies, Belgian Malinois and German Shepherds use tactics of hunter and prey to intimidate and keep control of herds and flocks. Their natural instinct to bring down an animal under their charge is muted by training. Other members of the group, including Welsh Corgis, Canaan dogs, and Cattle dogs herd with a more aggressive demeanor and make use of body design to elude the defences of their charges.
* Gun dog breeds used in hunting—that is, pointers, setters, spaniels, and retrievers—have an intermediate degree of paedomorphism; they are at the point where they share in the pack's hunting behavior, but are still in a junior role, not participating in the actual attack. They identify potential prey and freeze into immobility, for instance, but refrain from then stalking the prey as an adult predator would do next; this results in the "pointing" behavior for which such dogs are bred. Similarly, they seize dead or wounded prey and bring it back to the "pack", even though they did not attack it themselves, that is, "retrieving" behavior. Their physical characteristics are closer to that of the mature wild canine than the sheepdog breeds, but they typically do not have erect ears, etc.

Intentionally cross-bred dogs, such as this poodle hybrid, have become so popular in contemporary times so as to be labeled "designer dogs".
Intentionally cross-bred dogs, such as this poodle hybrid, have become so popular in contemporary times so as to be labeled "designer dogs".

* Scenthounds maintain an intermediate body type and behavior pattern that causes them to actually pursue prey by tracking their scent, but tend to refrain from actual individual attacks in favor of vocally summoning the pack leaders (in this case, humans) to do the job. They often have a characteristic vocalization called a bay. Some examples are the Beagle, Bloodhound, Basset Hound, Coonhound, Dachshund, Fox Hound, Otter Hound, and Harrier.
* Sighthounds, who pursue and attack perceived prey on sight, maintain the mature canine size and some features, such as narrow chest and lean bodies, but have largely lost the erect ears of the wolf and thick double layered coats. Some examples are the Afghan, Borzoi, Saluki, Sloughi, Pharaoh Hound, Azawakh, Whippet, and Greyhound.
* Mastiff-types are large dogs, both tall and massive with barrel-like chests, large bones, and thick skulls. They have traditionally been bred for war, protection, and guardian work.
* Bulldog-types are medium sized dogs bred for combat against both wild and domesticated animals. These dogs have a massive, square skull and large bones with an extremely muscular build and broad shoulders.
* Terriers similarly have adult aggressive behavior, famously coupled with a lack of juvenile submission, and display correspondingly adult physical features such as erect ears, although many breeds have also been selected for size and sometimes dwarfed legs to enable them to pursue prey in their burrows.

The least paedomorphic behavior pattern may be that of the basenji, bred in Africa to hunt alongside humans almost on a peer basis; this breed is often described as highly independent, neither needing nor appreciating a great deal of human attention or nurturing, often described as "catlike" in its behavior. It too has the body plan of an adult canine predator. Of course, dogs in general possess a significant ability to modify their behavior according to experience, including adapting to the behavior of their "pack leaders"—again, humans. This allows them to be trained to behave in a way that is not specifically the most natural to their breed; nevertheless, the accumulated experience of thousands of years shows that some combinations of nature and nurture are quite daunting, for instance, training whippets to guard flocks of sheep.

Breed popularity

Breed popularity varies widely over time[11] and in different parts of the world and different segments of the population. Counting by American Kennel Club (AKC) registration (not by licensing registration or by United Kennel Club (UKC) registration, which could present different statistics), the Labrador Retriever has been the United States's most commonly registered breed of dog since 1991. [12] However, even within parts of the United States, popularity varies; for example, in 2005 the most-registered breed in New York City was the Poodle while the Yorkshire Terrier was the second-most-registered breed in Houston. [13] However, animal shelters in many parts of the United States report that the most-commonly available dog for adoption is the American Pit Bull Terrier or pit bull-type mixes, making up as much as 20% of dogs available for adoption, none of which would be registered with the AKC.[14] Two decades ago, in 1983, the AKC's top two registered breeds were the American Cocker Spaniel and the Poodle.[15]

In 2006 a British dog journal published an exhaustive research paper which pooled together quantifiable dog breed facts such as average lifespan, cost of maintenance, insurance claims made, known behavioral faults and various other criteria in order to arrive at the conclusion that the Staffordshire Bull Terrier is the 'highest scoring' dog breed in terms of desirable and established pedigree dog breed traits. [16]

In the United Kingdom, The Kennel Club reports that the most-registered breed from at least 1999 to 2005 was the Labrador Retriever. It rounds out the top three for 1999 to 2005 with the German Shepherd Dog, also popular in the States, and the English Cocker Spaniel[17] , which is no longer even in the top ten in the States. In the United Kindom a national dog adoption and rescue service indicates that the most common breed appearing in shelters is the Greyhound followed by the Staffordshire Bull Terrier. [18]

Physical characteristics

Main article: Dog anatomy

Many dogs, such as this American Water Spaniel, have had their natural hunting instincts suppressed or altered to suit human needs.
Many dogs, such as this American Water Spaniel, have had their natural hunting instincts suppressed or altered to suit human needs.

See also: Dog health

Modern dog breeds show more variation in size, appearance, and behavior than any other domestic animal. Within the range of extremes, dogs generally share attributes with their wild ancestors, the wolves. Dogs are predators and scavengers, possessing sharp teeth and strong jaws for attacking, holding, and tearing their food. Although selective breeding has changed the appearance of many breeds, all dogs retain basic traits from their distant ancestors. Like many other predatory mammals, the dog has powerful muscles, fused wristbones, a cardiovascular system that supports both sprinting and endurance, and teeth for catching and tearing. Compared to the bone structure of the human foot, dogs technically walk on their toes.

Sight

Like most mammals, dogs are dichromats and have color vision equivalent to red-green color blindness in humans.[19][20]

Different breeds of dogs have different eye shapes and dimensions, and they also have different retina configurations.[21] Dogs with long noses have a "visual streak" which runs across the width of the retina and gives them a very wide field of excellent vision, while those with short noses have an "area centralis" — a central patch with up to three times the density of nerve endings as the visual streak — giving them detailed sight much more like a human's.

Some breeds, particularly the sighthounds, have a field of vision up to 270° (compared to 180° for humans), although broad-headed breeds with short noses have a much narrower field of vision, as low as 180°.[19][20]

Hearing

Dogs detect sounds as low as the 16 to 20 Hz frequency range (compared to 20 to 70 Hz for humans) and above 45 kHz[22] (compared to 13 to 20 kHz for humans),[20] and in addition have a degree of ear mobility that helps them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate and raise or lower a dog's ear. Additionally, a dog can identify a sound's location much faster than a human can, as well as hear sounds up to four times the distance that humans are able to. Those with more natural ear shapes, like those of wild canids like the fox, generally hear better than those with the floppier ears of many domesticated species.

Smell
Scent hounds, especially the Bloodhound, are iconic for their keen sense of smell
Scent hounds, especially the Bloodhound, are iconic for their keen sense of smell

Dogs have nearly 220 million smell-sensitive cells over an area about the size of a pocket handkerchief (compared to 5 million over an area the size of a postage stamp for humans). Some breeds have been selectively bred for excellence in detecting scents, even compared to their canine brethren. What information a dog actually detects when he is scenting is not perfectly understood; although once a matter of debate, it now seems to be well established that dogs can distinguish two different types of scents when trailing, an air scent from some person or thing that has recently passed by, as well as a ground scent that remains detectable for a much longer period. The characteristics and behavior of these two types of scent trail would seem, after some thought, to be quite different, the air scent being intermittent but perhaps less obscured by competing scents, whereas the ground scent would be relatively permanent with respect to careful and repetitive search by the dog, but would seem to be much more contaminated with other scents. In any event, it is established by those who train tracking dogs that it is impossible to teach the dog how to track any better than it does naturally; the object instead is to motivate it properly, and teach it to maintain focus on a single track and ignore any others that might otherwise seem of greater interest to an untrained dog. An intensive search for a scent, for instance searching a ship for contraband, can actually be very fatiguing for a dog, and the dog must be motivated to continue this hard work for a long period of time.

Coat color

Domestic dogs often display the remnants of counter-shading, a common natural camouflage pattern. The general theory of countershading is that an animal that is lit from above will appear lighter on its upper half and darker on its lower half where it will usually be in its own shade. This is a pattern that predators can learn to watch for. A countershaded animal will have dark coloring on its upper surfaces and light coloring below. This reduces the general visibility of the animal. One reminder of this pattern is that many breeds will have the occasional "blaze", stripe, or "star" of white fur on their chest or undersides.

Sprint metabolism

Dogs can generate large amounts of energy for a short period of time. A dog's heart and lungs are oversized relative to its body and its normal everyday needs. A dog also has relatively more red blood cells than a human. Most of the time the dog will keep the extra red blood cells stored in its spleen. When the animal enters into a situation where its full metabolism is required, such as play, catching game, or fighting other dogs, the extra cells are released into the bloodstream. The "oversized" heart and lungs will now be running at full capacity, and the animal will have an enhanced ability to engage in aerobic activity. This activity will produce internal heating. Dogs, being covered in fur, are limited in their ability to cool down. After a short time the animal must either cease its athletic activity or risk harming itself from overheating. One can easily observe this pattern of intense activity followed by rest periods in puppies. During the rest phase the spleen collects red blood cells and the animal may pant to cool down.

Behavior and Intelligence

Further information: Category:Dog training and behavior

Dogs are very social animals, but their personality and behavior vary with breed as well as how they are treated by their owners and others who come in contact with them. It is not uncommon for dogs to attack humans and other animals; however, this is usually because of lack of care or improper upbringing by its owner.

Intelligence

Main article: Dog intelligence

Many dogs can be trained to skillfully perform tasks not natural to canines, such as this Golden Retriever in a dog agility competition.
Many dogs can be trained to skillfully perform tasks not natural to canines, such as this Golden Retriever in a dog agility competition.

Dogs are valued for their intelligence. This intelligence is expressed differently with different breeds and individuals, however. For example, Border Collies are noted for their ability to learn commands, while other breeds may not be so motivated towards obedience, but instead show their cleverness in devising ways to steal food or escape from a yard. Being highly adaptable animals themselves, dogs have learned to do many jobs as required by humans over the generations. Dogs are employed in various roles across the globe, proving invaluable assets in areas such as search-and-rescue; law enforcement (including attack dogs, sniffer dogs and tracking dogs); guards for livestock, people or property; herding; Arctic exploration sled-pullers; guiding the blind and acting as a pair of ears for the deaf; assisting with hunting, and a great many other roles which they may be trained to assume. Most dogs rarely have to deal with complex tasks and are unlikely to learn relatively complicated activities (such as opening doors) unaided. Some dogs (such as guide dogs for the visually impaired) are specially trained to recognize and avoid dangerous situations.

Evaluation of a dog's intelligence

The meaning of "intelligence" in general, not only in reference to dogs, is hard to define. Some tests measure problem-solving abilities and others test the ability to learn in comparison to others of the same age. Defining it for dogs is just as difficult. It is likely that dogs do not have the ability to premeditate an action to solve a problem.

For example, the ability to learn quickly could be a sign of intelligence. Conversely it could be interpreted as a sign of blind subservience and a desire to please. In contrast, some dogs who do not learn very quickly may have other talents. An example is breeds that are not particularly interested in pleasing their owners, such as Siberian Huskies. Huskies are often fascinated with the myriad of possibilities for escaping from yards, catching small animals, and often figuring out on their own numerous inventive ways of doing both.

Assistance dogs are also required to be obedient at all times. This means they must learn a tremendous number of commands, understand how to act in a large variety of situations, and recognize threats to their human companion, some of which they might never before have encountered.

Many owners of livestock guardian breeds believe that breeds like the Great Pyrenees or the Kuvasz are not easily trained because their stubborn nature prevents them from seeing the point of such commands as “sit” or “down”. Hounds may also suffer from this type of ranking. These dogs are bred to have more of a "pack" mentality with other dogs and less reliance on a master's direct commands. While they may not have the same kind of intelligence as a Border Collie, they were not bred to learn and obey commands quickly, but to think for themselves while trailing game.

Human relationships

Dogs are highly social animals sometimes regarded as the most social animals on earth. This can account for their trainability, playfulness, and ability to fit into human households and social situations. This similarity has earned dogs a unique position in the realm of interspecies relationships despite being one of the most effective and potentially dangerous predators. Dogs and humans at times co-operate in some of the most effective hunting in the animal world; in that context, dogs are superpredators.

The loyalty and devotion that dogs demonstrate as part of their natural instincts as pack animals closely mimics the human idea of love and friendship, leading many dog owners to view their pets as full-fledged family members. Conversely, dogs seem to view their human companions as members of their pack, and make few, if any, distinctions between their owners and fellow dogs. Dogs fill a variety of roles in human society and are often trained as working dogs. For dogs that do not have traditional jobs, a wide range of dog sports provide the opportunity to exhibit their natural skills. In many countries, the most common and perhaps most important role of dogs is as companions.
Dogs have been used for a wide variety of purposes in human history.
Dogs have been used for a wide variety of purposes in human history.

Dogs have lived and worked with humans in so many roles that their loyalty has earned them the unique sobriquet "man's best friend".[23] However, some cultures consider dogs to be unclean. In some parts of the world, dogs are raised as livestock to produce dog meat for human consumption. In many places, consumption of dog meat is discouraged by social convention or cultural taboo.

Dog communication

Main article: Dog communication

Laughter in dogs

Laughter might not be confined or unique to humans, despite Aristotle's observation that "only the human animal laughs". The differences between chimpanzee and human laughter may be the result of adaptations that have evolved to enable human speech. However, some behavioral psychologists argue that self-awareness of one's situation, or the ability to identify with somebody else's predicament, are prerequisites for laughter, so animals are not really laughing in the same way that humans do. The dog-laugh sounds similar to a normal pant. However by analyzing the pant using a spectrograph, this pant varies with bursts of frequencies, resulting in a laugh. When this recorded dog-laugh vocalization is played to dogs in a shelter setting, it can initiate play, promote pro-social behavior, and decrease stress levels. In a study by Simonet, Versteeg, and Storie, one hundred and twenty subject dogs residing in a mid-size county animal shelter were observed. Dogs ranging from 4 months to 10 years of age were compared with and without exposure to a dog-laugh recording. The stress behaviors measured included panting, growling, salivating, pacing, barking, cowering, lunging, play-bows, sitting, orienting and lying down. The study resulted in positive findings. Exposure to the dog laughing recording resulted in the following: significantly reduced stress behaviors, increased tail wagging and the display of a play-face when playing was initiated, and more frequent pro-social behavior such as approaching and lip licking. This research suggests exposure to dog-laugh vocalizations can calm and possibly increase shelter adoptions. (Simonet, Versteeg, & Storie 2005) A dog laughter sample: Simonet 2005.

Reproduction

Life cycle

Main article: Canine reproduction

A Catahoula Leopard mother nursing her litter of puppies.
A Catahoula Leopard mother nursing her litter of puppies.

In domestic dogs, sexual maturity (puberty) begins to happen around age 6 to 12 months for both males and females, although this can be delayed until up to two years old for some large breeds. Adolescence for most domestic dogs is around 12 to 15 months, beyond which they are for the most part more adult than puppy. As with other domesticated species, domestication has selectively bred for higher libido and earlier and more frequent breeding cycles in dogs, than in their wild ancestors. Dogs remain reproductively active until old age

Most female dogs have their first estrous cycle between 6 and 12 months, although some larger breeds delay until as late as 2 years. Females experience estrous cycles biannually ,during which her body prepares for pregnancy, and at the peak she will come into estrus, during which time she will be mentally and physically receptive to copulation.

Dogs bear their litters roughly 56 to 72 days after fertilization, although the length of gestation can vary from. An average litter consists of about six puppies, though this number may vary widely based on the breed of dog. Toy dogs generally produce from one to four puppies in each litter, while much larger breeds may average as many as 12 pups in each litter.

Spaying and neutering

For more details on this topic, see Spaying and neutering.

Spaying (females only) and neutering (both genders but more usually males) refers to the sterilization of animals, usually by removal of the male's testicles or the female's ovaries and uterus, in order to eliminate the ability to procreate, and reduce sex drive. Neutering has also been known to reduce aggression in male dogs, but has been shown to occasionally increase aggression in female dogs.[24]

Animal control agencies in the United States and the ASPCA advise that dogs not intended for further breeding should be spayed or neutered so that they do not have undesired puppies.[25]

Because of the overpopulation of dogs in some countries, puppies born to strays or as the result of accidental breedings often end up being killed in animal shelters. Spaying and neutering can also decrease the risk of hormone-driven diseases such as mammary cancer, as well as undesired hormone-driven behaviors. However, certain medical problems are more likely after neutering, such as urinary incontinence in females[26] and prostate cancer in males.[27] The hormonal changes involved with sterilization are likely to somewhat change the animal's personality, however, and some object to spaying and neutering as the sterilization could be carried out without the excision of organs.

It is not essential for a female dog to either experience a heat cycle or have puppies before spaying, and likewise, a male dog does not need the experience of mating before neutering.

Female cats and dogs are seven times more likely to develop mammary tumors if they are not spayed before their first heat cycle. [28] The high dietary estrogen content of the average commercial pet food as well as the estrogenic activity of topical pesticides [citation needed] may be contributing factors in the development of mammary cancer, especially when these exogenous sources are added to those normal estrogens produced by the body. Dog food containing soybeans or soybean fractions have been found to contain phytoestrogens in levels that could have biological effects when ingested longterm.[29]

Gender-preservative surgeries such as vasectomy and tubal ligation are possible, but do not appear to be popular due to the continuation of gender-specific behaviors and disease risks.

Overpopulation
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According to the Humane Society of the United States, 3-4 million dogs and cats are euthanized each year in the United States and many more are confined to cages in shelters because there are many more animals than there are homes. Spaying or neutering dogs helps keep overpopulation down.[30] Local humane societies, SPCAs and other animal protection organizations urge people to spay or neuter their pets and to adopt animals from shelters instead of purchasing them. Several notable public figures have spoken out against animal over population, including Bob Barker. On his game show, The Price is Right, Barker stressed the issue at the end of every episode, saying: "Help control the pet population. Have your pets spayed or neutered."

Working, utility and assistance dogs

Main article: Working dog

There are several types of working dogs:
Mixed-breed dogs, such as this Australian border collie cross, are often desired as working animals for their robust health and intelligence.
Mixed-breed dogs, such as this Australian border collie cross, are often desired as working animals for their robust health and intelligence.

* Assistance dogs which help the seeing and hearing-impaired. Others are trained to help those with epilepsy and psychiatric disorders, by detecting the onset of the condition so they can seek help. The typical assistance dog is a sociable breed such as a Labrador Retriever.
* The detection dog, which is a dog trained to and works at using its senses (almost always the sense of smell) to detect substances such as explosives or illegal drugs.
* Guard dogs trained to protect property. These are commonly Doberman Pinschers, German Shepherds and Rottweilers or mixes of these breeds.
* Herding dogs trained to protect livestock in the range against predators. Sheepdogs are a common breed.
* Hunting dogs that search for and retrieve game. Pointers and retrievers are typical hunting breeds.
* Police dogs, typically German Shepherds, are trained to assist law enforcement officers.
* Rescue dogs that search for missing humans. The archetypal breeds are the St. Bernard and the Bloodhound, although any dog with a good sense of smell can be trained to perform this task.
* Therapy dogs, with friendly and gentle temperaments, trained to provide comfort and affection to hospitalized and institutionalized patients.
* The war dog, used by the military to detect mines and enemy soldiers.

Show and sport (competition) dogs
German Shepherd Dogs at the Westminster Kennel Club dog show
German Shepherd Dogs at the Westminster Kennel Club dog show

Main article: Show dog
See also: Dog sport

Owners of dogs often enter them in competitions, whether show (breed conformation shows) or sports, including dog racing & dog sledding. The winners garner much prestige and prize money. These dogs are often bred specifically for competition, which may not be entirely beneficial for the breeds due to the magnification of hereditary defects.

* Dog agility is a sport in which dogs complete a timed obstacle course.
* Dogsled racing is a winter sport where a team of dogs, usually high performance mixed breed dogs called Alaskan Huskies or Eurohounds, pull a sled and driver (called a musher). The Iditarod Trail Sled Dog Race is perhaps the most famous of these races. Dog sledding is an ancient form of transportation and still a very effective way of moving freight across this type of terrain.
* Dog racing, almost always Greyhounds, involves dogs racing at betting tracks in a sport not unlike horseracing, reaching speeds of 40 miles per hour. Elsewhere, Dachshunds are often raced, as a humorous sidelight in charity events.
* Dog fighting and dog baiting are blood sports involving dogs. They are illegal in most jurisdictions, but are still performed underground. In some areas, the illegal practice is thriving[31].
* The show dog, purebreds entered in conformation dog shows and evaluated by how closely they match the ideal characteristics of the breed. The Westminster Kennel Club Dog Show is one of the most famous of this type of competition.

Dog health

Main article: Dog health
Further information: Category:Dog health

Morbidity (Illness)

Dogs are susceptible to various diseases, ailments, and poisons, some of which affect humans in the same way, others of which are unique to dogs. Dogs, like all mammals, are also susceptible to heat exhaustion when dealing with high levels of humidity and/or extreme temperatures.[32]

Diseases

Infectious diseases commonly associated with dogs include rabies (hydrophobia), canine parvovirus, and canine distemper. Inherited diseases of dogs can include a wide range from hip dysplasia and medial patellar luxation to epilepsy and pulmonic stenosis. Canines can get just about anything a human can get (excluding many infections which are species specific) like hypothyroidism, cancer, dental disease, heart disease, etc.

Two serious medical conditions affecting dogs are pyometra, affecting unspayed females of all types and ages, and bloat, which affects the larger breeds or deep chested dogs. Both of these are acute conditions, and can kill rapidly; owners of dogs which may be at risk should learn about such conditions as part of good animal care.

See also: Vaccination of dogs

First generation hybrids (such as this terrier mix) often are healthier than either parent due to the genetic phenomenon of heterosis or "hybrid vigor".
First generation hybrids (such as this terrier mix) often are healthier than either parent due to the genetic phenomenon of heterosis or "hybrid vigor".

Parasites

Common external parasites are various species of fleas, ticks, and mites. Internal parasites include hookworms, tapeworms, roundworms, and heartworms. See also CVBD (Canine Vector-Borne Diseases).

Common physical disorders

Some breeds of dogs are also prone to certain genetic ailments, such as hip dysplasia, luxating patellas, cleft palate, blindness, or deafness. Dogs are also susceptible to the same ailments that humans are, including diabetes, epilepsy, cancer, and arthritis. Gastric torsion and bloat is a dangerous problem in some large-chested breeds.

Mortality (Lifespan and causes of death)

Main articles: Dog years and Aging in dogs

The typical lifespan of dogs varies widely among breeds. Based on questionnaire surveys of owners in the UK, Denmark, and the USA/Canada, the median longevity of most dog breeds is between 10 and 13 years.[33][34][35][36] The breed with the dubious distinction of the shortest lifespan (among breeds for which there is a questionnaire survey with a reasonable sample size) is the Dogue de Bordeaux with a median longevity of about 5.2 years, but several breeds, including Miniature Bull Terrier, Bulldog, Nova Scotia Duck Tolling Retriever, Bloodhound, Irish Wolfhound, Greater Swiss Mountain Dog, Great Dane, and Mastiff, are nearly as short-lived, with median longevities between 6 and 7 years.[36] On the other end of the spectrum, the longest-lived breeds, including Toy Poodle, Border Terrier, Miniature Dachshund, Miniature Poodle, and Tibetan Spaniel, have median longevities between 14 and 15 years.[36] The median longevity of mixed breed dogs (average of all sizes) is one or more years longer than that of purebred dogs (all breeds averaged).[35][34][36][37] As a rule of thumb, small breeds are longer-lived than large breeds, but some of the longest lived large breeds have median longevities nearly as large as those of the shortest lived small breeds, and some of the breeds with the shortest longevities are medium-sized.[38]

"Median longevity" refers to the age at which half the dogs in a population have died and half are still alive. Individual dogs, even in breeds with low median longevities, may live well beyond the median. The dog widely reported to be the longest-lived on record is "Bluey," purportedly born in 1910 in Australia. He died in 1939 at the age of 29.5 years. Bluey is usually identified as an Australian Cattle Dog, but the first Australian Cattle Dog breed standard was written in 1902,[39] only eight years before Bluey's birth. It is unclear how closely Bluey was related to the breed as it exists today. The Bluey record is anecdotal and unverified.[40] The longest verified records are of dogs living to 24 years.[40]

Diet

Nutrition
There are a great quantity of commercial foods and treats marketed for dogs, and not all are recommended as part of a balanced, healthy diet.
There are a great quantity of commercial foods and treats marketed for dogs, and not all are recommended as part of a balanced, healthy diet.

At present, there is some debate as to whether domestic dogs should be classified as omnivores or carnivores, by diet. The classification in the Order Carnivora does not necessarily mean that a dog's diet must be restricted to meat; unlike an obligate carnivore, such as the cat family with its shorter small intestine, a dog is dependent on neither meat-specific protein nor a very high level of protein in order to fulfill its basic dietary requirements. Dogs are able to healthily digest a variety of foods including vegetables and grains, and in fact dogs can consume a large proportion of these in their diet. Wild canines not only eat available plants to obtain essential amino acids, but also obtain nutrients from vegetable matter from the stomach and intestinal contents of their herbivorous prey, which they usually consume. Domestic dogs can survive healthily on a reasonable and carefully designed vegetarian diet, particularly if eggs and milk products are included. Some sources suggest that a dog fed on a strict vegetarian diet may develop dilated cardiomyopathy since it lacks L-carnitine,[41] however, maintaining a balanced diet is also a factor since L-carnitine is found naturally in many nuts, seeds, beans, vegetables, fruits and whole grains. In the wild, dogs can survive on a vegetarian diet when animal prey is not available. However it has been noted, both by observation of extremely stressful conditions such as the Iditarod Trail Sled Dog Race and by scientific studies of similar conditions, that high-protein (approximately 40%) diets including meat help prevent damage to muscle tissue. (This research is also true for some other mammals.) This level of protein corresponds to the percentage of protein found in the wild dog's diet when prey is abundant; higher levels of protein seem to confer no added benefit.

Dogs frequently eat grass, which is a harmless activity. Explanations abound, but rationales such as that it neutralizes acid, or that eating grass might make the dog vomit, so dogs eat grass to remove unwanted substances from their stomachs, are at best educated guesses. Indeed, dogs may eat grass when they have an upset stomach in order to cause vomiting. The grass binds with the offending material. Then the grass acts as an irritant and causes the dog to vomit, bringing up the poisonous material at the same time..[42] Dogs do vomit more readily than humans, as part of their typical feeding behavior of gulping down food then regurgitating indigestible bones, fur, etc. This behavior is typical of pack feeding in the wild, where the most important thing is to get as much of the kill as possible before others consume it all. Individual domestic dogs, however, may be very "picky" eaters, in the absence of this evolutionary pressure.

Also, dogs eat grass, because it's a normal part of their diet. Wild canine relatives of dogs — wolves and foxes — eat all of an animal when they catch it. They end up, therefore, eating grasses and plants in the intestines of these animals. Domestic dogs, eating prepared dog food, lack this part of their diet and so they eat grass. Finally, dogs may appear to eat grass, when they are just running the blades through their mouth to gather information. Their sense of smell and taste may act together to detect if other animals have walked through their area or urinated on the grass.[43]

Dangerous substances

Some foods commonly enjoyed by humans are dangerous to dogs, including chocolate (Theobromine poisoning), onions, grapes and raisins,[44] some types of gum, certain sweeteners[45] and Macadamia nuts. It is currently believed that the only dangerous substance in chocolate is cocoa, meaning that forms of chocolate without this ingredient, such as white chocolate may become re-evaluated for their safety in the future.

The acute danger from grapes and raisins has been uncovered only since about 2000, and made public slowly since then. At present the cause is not known. Whatever the reason, since only small quantities are necessary to induce acute renal failure, dogs should not be fed grapes or raisins, and sultanas and currants should likely be withheld as well.
Despite the popular notion, bones are not the ideal treat for dogs.
Despite the popular notion, bones are not the ideal treat for dogs.

Cooked bones should never be given to dogs, as the heat changes the chemical and physical properties so that they cannot be chewed properly, splintering into jagged shards, and resist digestion.

Human medications should not be given to a dog as a substitute for their regular medication as some can be especially toxic, especially paracetamol/acetaminophen (Tylenol). Alcoholic beverages pose much of the same hazards to dogs as they do to humans.

Dogs may also find some poisons attractive, including antifreeze, snail bait, slug bait, insect bait, and rodent poisons. Antifreeze may be one of the most insidious of poisons to dogs because of its sweet taste and because a dog may walk upon or lie down upon a spill of it or its residue and then lick it off. Dogs must be kept strictly away from antifreeze and not allowed access to any place that has had a spill of it that has not been completely removed.

Plants such as caladium, dieffenbachia and philodendron will cause throat irritations that will burn the throat going down as well as coming up. Hops are particularly dangerous and even small quantities can lead to malignant hyperthermia.[46]

Amaryllis, daffodil, english ivy, iris, and tulip (especially the bulbs) cause gastric irritation and sometimes central nervous system excitement followed by coma, and, in severe cases, even death.

Ingesting foxglove, lily of the valley, larkspur and oleander can be life threatening because the cardiovascular system is affected. Equally life threatening is the yew which affects the nervous system. If any of these plants are ingested, get the dog to a veterinarian immediately.

Many household cleaners such as ammonia, bleach, disinfectants, drain cleaner, soaps, detergents, and other cleaners, mothballs and matches are dangerous to dogs, as are cosmetics such as deodorants, hair coloring, nail polish and remover, home permanent lotion, and suntan lotion.

Zinc toxicity, mostly in the form of the ingestion of US pennies minted after 1982, is commonly fatal in dogs where it causes a severe hemolytic anemia.[47]

Dogs will occasionally eat their own feces or the feces of other species if available, such as that of cats and horses. This is known as coprophagia, and may be indicative of boredom or hunger, although there is no definitive reason known. Owners of both cats and dogs may find that dogs treat the catbox as a snack bar. Such behavior should be discouraged, as it could lead to Toxoplasmosis. Dogs seem to have different preferences in relation to eating feces. Some are attracted to the stools of deer, cows, or horses. Others will eat the stools of other dogs. And a great many dogs are attracted to cat droppings, possibly because cat foods are very high in protein and the dogs are going after undigested nutrients.[48]

Dog abuse

Main article: Cruelty to animals

Cruelty to dogs refers to treatment that causes unacceptable suffering or harm. What qualifies as unacceptable suffering varies among countries and cultures. Cruelty can be passive, typified by simple neglect, or active, with malicious intent.[49]

Malicious treatment of a dog can lead to dog attacks upon not only the abuser but also innocent people.

Signs of dog abuse

Signs of dog abuse include:

* Unusually frightened, fearful or subdued
* Fractures
* Bruising
* Eye injuries
* Scalds and burns
* Signs of malnutrition
* Significant matting or other poor grooming indicators
* Ignored health problems
* Injury history incompatible with injury or owner refusing to comment on how injury occurred
* Owner showing lack of concern for animal’s injuries
* Owner exhibiting Munchausen’s Syndrome by Proxy (MSP)

Dog meat for human consumption

Main article: Dog meat

In some countries, certain dog breeds, apart from being kept as pets, are raised on farms and slaughtered for consumption. In countries where dogs are particularly popular as household pets, consumption of dogs is generally taboo and considered an abhorrent cultural practice, but there are exceptions, such as Vietnam, where dogs are popular as both pets and meat.[50]

Although the Chinese were the first to domesticate the dog and keep them as pets, dog meat has been a source of food in China from at least the time of Confucius, and possibly even before. Ancient writings from the Zhou Dynasty referred to the 'three beasts' (which were bred for food), including pig, goat, and dog. Mencius, the philosopher, recommended dog as the tastiest of all meats.

Dogs in religion
Statue of Saint Roch with his dog, in Prague, Czech Republic
Statue of Saint Roch with his dog, in Prague, Czech Republic

Chinese tradition

The dog is one of the 12 animals honored in Chinese astrology. The second day of the Chinese New Year is considered to be the birthday of all dogs and Chinese people often take care to be kind to dogs on that day.

Christianity

Jesus told the story of the poor man Lazarus, whose sores were licked by street dogs. This has traditionally been seen as showing Lazarus's wretched situation. However, some modern commentators have pointed out that the dogs' saliva could have beneficial effects on the sores and that this could be seen as an example of the dogs' unconditional love.

The Catholic Church recognizes Saint Roch (also called Saint Rocco), who lived in the early 1300s in France, as the patron saint of dogs. It is said that he caught the black plague while doing charitable work and went into the forest expecting to die. There he was befriended by a dog which licked his sores and brought him food and he was able to recover. The feast day of Saint Roch is August 16. [51]

Islam

See also Islam and animals

Islamic tradition considers dogs to be unclean and most Muslims do not keep pet dogs. There are a number of traditions concerning Muhammad's attitude towards dogs. He said that the company of dogs, except as helpers in hunting, herding, and home protection, voided a portion of a Muslim's good deeds. On the other hand, he advocated kindness to dogs and other animals. It is said that he told a prostitute that if she were to give water to a thirsty dog her sins would be forgiven.

Atheism and criticism of religion

In an article in the New York Times Magazine atheist Natalie Angier quoted Frans de Waal, a primatologist at Emory University:

"I've argued that many of what philosophers call moral sentiments can be seen in other species. In chimpanzees and other animals, you see examples of sympathy, empathy, reciprocity, a willingness to follow social rules. Dogs are a good example of a species that have and obey social rules; that's why we like them so much, even though they're large carnivores."[52]

A Newfoundland, the breed Byron eulogized, painted by Edwin Henry Landseer, 1802-1873
A Newfoundland, the breed Byron eulogized, painted by Edwin Henry Landseer, 1802-1873

In 1808 the English poet Byron expressed similar thoughts in his famous poem Epitaph To a Dog:

But the poor dog, in life the firmest friend,
The first to welcome, foremost to defend,
Whose honest heart is still his master’s own,
Who labors, fights, lives, breathes for him alone,
Unhonored falls, unnoticed all his worth,
Denied in heaven the soul he held on earth –
While man, vain insect! hopes to be forgiven,
And claims himself a sole exclusive heaven.[53]

See also

* List of dog breeds
* List of most popular dog breeds
* List of famous dogs

Other:

* Dog paddle - basic swimming stroke
* Bark (dog)
* Dog licence
* Dog odor
* Fear of dogs
* Wolf-dog hybrid
* The Dog king - Scandinavian tradition

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